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Fire is an essential component of ecosystems and acts as key driver of biogeochemical cycling with impacts on vegetation structure and composition, soil conditions, and climate feedbacks. Fire behavior and effects vary based on the types of fuel, fuel dryness, and frequency of ignition. Due to changing climate and land use patterns, fire danger is increasing in many regions globally, and fires are having increasingly devastating impacts on human health, infrastructure, and ecosystem services. Recurrent fires help to determine the distribution of trees and grasses, and overall fuel load, which inform the behavior of future fires. Process-based models can be used to capture multi-scale impacts at the forest stand-level up to the landscape level, and across minutes to centuries, but must capture variation in fire behavior and intensity as a function of the fuel and climate, from high intensity forest crown fires within boreal regions to low intensity rapid grass fires of the tropics. Fire model development demonstrates our ability to capture large scale fire influenced biogeography and vegetation distribution at the earth system scale through vegetation traits and fire feedbacks. At the individual stand scale the importance of interactions and feedbacks between above and below ground process is essential to capturing the vegetation dynamics across boreal forest systems. Improving the mechanics of including plant physiology and specifically live fuel moisture content is the next step to advancing the capability of process based models to inform fire research. Advances in the testing and creation of a mechanistic live fuel moisture model demonstrate the foundation for future live fuel dynamics research that can be informed by field and remote sensing information. Improved remote sensing, technological and modeling capabilities support a more comprehensive and cohesive fire response that will be better equipped to overcome current barriers and anticipate the new reality of fires in a warming world.