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How does drought affect residential water demand and price elasticity?

Urban water scarcity is an important social and economic concern, particularly as the intensity, duration, and frequency of droughts is increasing in many regions. We consider whether drought induces changes to water demand and the price elasticity of demand for water that may last beyond a drought’s official end date. If drought shocks prompt long-term changes in water demand behavior, and these changes occur at broad geographic scale, they could have important implications for modeling adaptive responses to water scarcity. We assemble a novel dataset on residential water demand and pricing in the western United States to test empirically for effects of drought on water demand and price elasticity. We perform our analysis with aggregate quantity, price, and drought data, accounting for endogenous prices under increasing-block water tariffs and using both average and marginal water fees in estimating water demand functions. Results are consistent with the hypothesis that households may become less price-sensitive after exposure to drought. However, we find no systematic evidence of long-run, drought-related reductions in water demand, itself.

demand hardening↗

Diverging Importance of Drought Stress for Maize and Winter Wheat in Europe

Understanding the drivers of yield levels under climate change is required to support adaptation planning and respond to changing production risks. This study uses an ensemble of crop models applied on a spatial grid to quantify the contributions of various climatic drivers to past yield variability in grain maize and winter wheat of European cropping systems (1984-2009) and drivers of climate change impacts to 2050. Results reveal that for the current genotypes and mix of irrigated and rainfed production, climate change would lead to yield losses for grain maize and gains for winter wheat. Across Europe, on average heat stress does not increase for either crop in rainfed systems, while drought stress intensifies for maize only. In low-yielding years, drought stress persists as the main driver of losses for both crops, with elevated CO2 offering no yield benefit in these years.

Europe↗

The Future of Evapotranspiration: Global Requirements for Ecosystem Functioning, Carbon and Climate Feedbacks, Agricultural Management, and Water Resources

The fate of the terrestrial biosphere is highly uncertain given recent and projected changes in climate. This is especially acute for impacts associated with changes in drought frequency and intensity on the distribution and timing of water availability. The development of effective adaptation strategies for these emerging threats to food and water security are compromised by limitations in our understanding of how natural and managed ecosystems are responding to changing hydrological and climatological regimes. This information gap is exacerbated by insufficient monitoring capabilities from local to global scales. Here, we describe how evapotranspiration (ET) represents the key variable in linking ecosystem functioning, carbon and climate feedbacks, agricultural management, and water resources, and highlight both the outstanding science and applications questions and the actions, especially from a space-based perspective, necessary to advance them.

Fisher, Joshua B.↗

Sensitivity Analysis of Drivers Water Shortage in the Los Angeles Region During Drought

The code and detailed step-by-step instructions for generating the model output data, processing results, and analysis and plotting are provided at https://github.com/IMMM-SFA/Ferencz_et_al_2026_ER_Water. The PyArtes model is a python adaptation of the Artes model. PyArtes uses many of the same input data and optimization model architecture as Artes. Documentation for the PyArtes model is provided in the Supplement to the paper. The primary data product are simulated monthly water shortages for indoor and outdoor demand under a large ensemble of drought scenarios (>13,000). The droughts are hypothetical and are not based on historical time series data of supply sources - though historical data did help inform ranges explored for supply parameters. Demands are informed by recent 2017-2021 water supply data. Demands used for the model can be accessed at https://github.com/IMMM-SFA/Ferencz_et_al_2026_ER_Water. Simulations resolve demand for over 90 water providers in the study region. The results report 36 months of water shortage data for each indoor and outdoor demand node. The study also developed a multilayer perceptron (MLP) neural network trained on a subset of the simulated shortage ensemble to emulate worst annual water shortage for a given set of parameter multipliers -- provided the parameter values fall within the ranges sampled in the ensemble. Emulated water shortages for synthetic ensembles are in the MLP-generated shortages folder. The MLP model was used to generate larger ensembles to support Sobol analysis that would have been extremely computationally expensive to simulate. Datasets provided in this repository*: Simulated shortages. These results are used for the analysis for Figures 5, 8, and 9 in the paper, and also to train the MLP emulator. .zip file containing outputs for the 13,312 scenario ensemble. Separate .csv files for indoor and outdoor shortage for each scenario. Rows = demand ids (~100), Columns = months (36) Units = acre-feet/month of shortage (shortage = monthly demand - supply). 1 acft = 1233.48 m^3 .csv files of aggregated shortages derived from the 13,312 ensemble Rows = scenarios (13,312), Columns = demand ids (~100) Units = acre-feet/year (either worst annual shortage or total shortage over the 3-year drought) .csv file of the parameter multipliers scenarios for the ensemble .csv file of the parameter ranges and baseline values the multipliers were applied to MLP-generated shortages. These results are used for Figures 4, 6, and 7 in the paper. mwd higher folder: scenario ensembles, emulated worst year total shortages (acft), and Sobol results Emulated shortages. Rows = scenarios, columns = demand ids, units acft Sobol results. Rows = demand ids, columns Sobol (S1, ST, or 95% confidence interval) value for each parameter mwd lower folder: scenario ensembles, emulated worst year total shortages (acft), and Sobol results same organization as mwd higher MLP performance: performance metrics (R^2, RMSE, BIAS, MAPE) for the testing subset (20% or 2,662 scenarios) and simulated vs emulated worst year shortage (acre-feet/year) for every demand node, MWD wholesale regions, and the entire study region (LAC). Supporting data for figures. Figure plotting scripts in the associated GitHub repo. These files support analysis and visualization. Geospatial Data used for plotting simulated water shortages and Sobol results. Dictionary of full names for demand nodes in the model and estimates of water supply by source type informed by Artes input files and California Urban Water Management Planning data: https://water.ca.gov/Programs/Water-Use-And-Efficiency/Urban-Water-Use-Efficiency/Urban-Water-Management-Plans *Readme files provided for each folder.

drought↗

Compound Heat and Moisture Extreme Impacts on Global Crop Yields Under Climate Change

Extreme heat, drought and moisture excess are increasingly co-occurring within a single growing season, impacting crop yields in global breadbasket regions. In this Review, we synthesize understanding of compound heat and moisture extremes, their impacts on global crop yields and implications for adaptation. Heat and moisture extremes and their impacts become compounded through crop-physiological interactions, heat–moisture couplings in the climate system and crop–atmosphere interactions. Since around 2000, these compound extremes, and hot droughts in particular, have been linked to especially poor harvests (up to 30% yield losses) in regions such as India, Ethiopia, the USA, Europe and Russia. However, in some cases, combinations of crop stresses might generate compensating effects. Compound extremes are projected to increase in frequency and amplitude in the future, but, owing to the biophysical interdependence among temperature, water and crop physiology, the net yield effects of such future compound extremes remain uncertain. Accordingly, compound extremes will necessitate comprehensive agricultural adaptation strategies geared towards multi-stress resilience, as adaptations that work for single climate stresses could be maladaptive under combined stresses. An integrated understanding of heat and water in soil–plant–atmosphere dynamics is urgently needed to understand risks and suitably adapt cropping systems to compounding climate impacts.

Agroecology↗

Adaptation and Response in Drylands (ARID): A Dryland Research Agenda and Campaign Strategy

Dryland ecosystems cover 41% of Earth’s land surfaces, account for 44% of cultivated lands and 60% of food sources, and make large contributions to the global water and carbon cycles. However, these ecosystems are experiencing unprecedented extremes including heatwaves, floods, and droughts as well as hotter temperatures and often declining water resource availability. These ecosystems are some of the most challenging to monitor given their high temporal variability with rapid response to their environment as well as vast spatial variability with intermixing of different plant species and life forms amongst bare soil coverage. The Adaptation and Response in Drylands (ARID) campaign was selected by National Aeronautics and Space Administration (NASA) as a scoping study to develop a research agenda for a dryland field campaign. Here, we detail our ARID science research agenda and implementation plan that were developed based on an extensive community engagement effort in over 160 events with over a thousand scientists, land managers, and Tribal communities between 2023 and 2024. The selected science themes cover drought and climate variability, ecosystem structure, function, and biodiversity, carbon cycle interannual variability and trends, and social ecological systems (land management and adaptation). We then detail our remote sensing, modeling, and field-based strategies to capture high temporal and high spatial resolution processes. Finally, our implementation strategy is presented which includes focus area selections in a core intensive western U.S. domain and distributed international domains. This strategy includes our overarching guiding principles of using multi-temporal airborne acquisitions and super sites as well as enhancing land management in co-development with end-user partners. While originally developed for NASA, our ARID report creates a blueprint for any future dryland field campaign, at any scale, that can be implemented widely for foundational and applied science objectives.

drylands↗

Two-Way Option Contracts That Facilitate Adaptive Water Reallocation in the Western United States

Many water markets in the western United States (U.S.) have the ability to reallocate water temporarily during drought, often as short-term water rights leases from lower value irrigated activities to higher value urban uses. Regulatory approval of water transfers, however, typically takes time and involves high transaction costs that arise from technical and legal analyses, discouraging short-term leasing. This leads municipalities to protect against drought-related shortfalls by purchasing large volumes of infrequently used permanent water rights. High transaction costs also result in municipal water rights rarely being leased back to irrigators in wet or normal years, reducing agricultural productivity. This research explores the development of a multi-year two-way option (TWO) contract that facilitates leasing from agricultural-to-urban users during drought and leasing from urban-to agricultural users during wet periods. The modeling framework developed to assess performance of the TWO contracts includes consideration of the hydrologic, engineered, and institutional systems governing the South Platte River Basin in Colorado where there is growing competition for water between municipalities (e.g., the city of Boulder) and irrigators. The modeling framework is built around StateMod, a network-based water allocation model used by state regulators to evaluate water rights allocations and potential rights transfers. Results suggest that the TWO contracts could allow municipalities to maintain supply reliability with significantly reduced rights holdings at lower cost, while increasing agricultural productivity in wet and normal years. Additionally, the TWO contracts provide irrigators with additional revenues via net payments of option fees from municipalities.

54 ENVIRONMENTAL SCIENCES↗

Adaptive Management Using Remote Sensing and Ecosystem Modeling in Response to Climate Variability and Invasive Aquatic Plants for the California Sacramento-San Joaquin Delta Water Resource

The California Sacramento-San Joaquin River Delta is the hub for California's water supply and supports important ecosystem services, agriculture, and communities in Northern to Southern California. Expansion of invasive aquatic plants in the Delta coupled with impacts of changing climate and long-term drought is detrimental to the San Francisco Bay/California Delta complex. NASA Ames Research Center and the USDA-ARS partnered with the State of California to develop science-based, adaptive-management strategies for invasive aquatic plant in the Sacramento-San Joaquin Delta. Specific mapping tools developed utilizing satellite and airborne platforms provide regular assessments of population dynamics on a landscape scale and support both strategic planning and operational decision making for resource managers. San Joaquin and Sacramento River watersheds water quality input to the Delta is modeled using the Soil-Water Assessment Tool (SWAT) and a modified SWAT tool has been customized to account for unique landscape and management of agricultural water supply and drainage within the Delta. Environmental response models for growth of invasive aquatic weeds are being parameterized and coupled with spatial distribution/biomass density mapping and water quality to study ecosystem response to climate and aquatic plant management practices. On the water validation and operational utilization of these tools by management agencies and how they are improving decision making, management effectiveness and efficiency will be discussed. The project combines science, operations, and economics related to integrated management scenarios for aquatic weeds to help land and water resource managers make science-informed decisions regarding management and outcomes.

Adaptive↗

EvoNet: A phylogenomic and systems biology approach to identify genes underlying plant survival in marginal, low‐N soils

The DOE‐BER “EvoNet” project investigates the genetic and molecular basis of plant resilience in extreme environments. We do this by identifying key genes that enable “extreme survivor” species to thrive in the nitrogen-poor soils of Chile’s hyper-arid Atacama Desert. Our collections focus on 32 Atacama extremophile species, including seven grass species with potential biofuel applications. To identify genes-of-importance to survival we compared genomic and transcriptomic profiles of extremophile species that thrive in the Atacama to those of closely related “sister” species from nitrogen-rich arid and mesic regions of California. Deep RNA sequencing and de novo transcriptome assembly across these triplet species sets supported a phylogenomic framework for identifying positively selected genes associated with adaptive divergence. Our integrative analysis combined ecological and environmental data, metagenomics, evolutionary and systems biology, and metabolomics. This enabled us to create an unprecedented framework for systematically understanding how non-model plants have adapted to survive in extreme conditions. Our resulting database of positively selected ortholog groups in the extremophile plants offers promising targets for engineering crop and biofuel species with enhanced resilience to drought and extreme weather. Additionally, our newest dataset explores and exploits a complementary metabolomic approach. This new aspect provides innovative strategies to manipulate plant cell metabolism, further supporting efforts to improve agricultural productivity in the face of extreme climates. Importantly, our combined evolutionary- and metabolomic-based strategies focused on convergent patterns of adaptation, providing a genetic and metabolomic toolkit for improving crop and biofuel resilience across diverse plant species. Finally, our novel exploration of ecological and evolutionary dynamics delivered to the community a phylogenomic computational pipeline called “PhyloGeneious.” Our continued adaptations of this pipeline are publicly available to expedite evolutionary genomic research for future scientific discoveries. In total, our DOE-BER has provided genomic, metabolomic, and computational strategies to understand how extremophile plants provide evolutionary and physiological targets for improving agricultural and biofuel production.

59 BASIC BIOLOGICAL SCIENCES↗

A Double Bootstrap Approach to Superposed Epoch Analysis to Evaluate Response Uncertainty

The association between climate variability and episodic events, such as the antecedent moisture conditions prior to wildfire or the cooling following volcanic eruptions, is commonly assessed using Superposed Epoch Analysis (SEA). In SEA the epochal response is typically calculated as the average climate conditions prior to and following all event years or their deviation from climatology. However, the magnitude and significance of the inferred climate association may be sensitive to the selection or omission of individual key years, potentially resulting in a biased assessment of the relationship between these events and climate. Here we describe and test a modified double-bootstrap SEA that generates multiple unique draws of the key years and evaluates the sign, magnitude, and significance of event-climate relationships within a probabilistic framework. This multiple resampling helps quantify multiple uncertainties inherent in conventional applications of SEA within dendrochronology and paleoclimatology. We demonstrate our modified SEA by evaluating the volcanic cooling signal in a Northern Hemisphere tree-ring temperature reconstruction and the link between drought and wildfire events in the western United States. Finally, we make our Matlab and R code available to be adapted for future SEA applications.

Dendroclimatology↗

Ground Water and Climate Change

As the world's largest distributed store of fresh water, ground water plays a central part in sustaining ecosystems and enabling human adaptation to climate variability and change. The strategic importance of ground water for global water and food security will probably intensify under climate change as more frequent and intense climate extremes (droughts and floods) increase variability in precipitation, soil moisture and surface water. Here we critically review recent research assessing the impacts of climate on ground water through natural and human-induced processes as well as through groundwater-driven feedbacks on the climate system. Furthermore, we examine the possible opportunities and challenges of using and sustaining groundwater resources in climate adaptation strategies, and highlight the lack of groundwater observations, which, at present, limits our understanding of the dynamic relationship between ground water and climate.

climate change↗

Climate Change Adaptation Science Activities at NASA Johnson Space Center

The Johnson Space Center (JSC), located in the southeast metropolitan region of Houston, TX is the prime NASA center for human spaceflight operations and astronaut training, but it also houses the unique collection of returned extraterrestrial samples, including lunar samples from the Apollo missions. The Center's location adjacent to Clear Lake and the Clear Creek watershed, an estuary of Galveston Bay, puts it at direct annual risk from hurricanes, but also from a number of other climate-related hazards including drought, floods, sea level rise, heat waves, and high wind events all assigned Threat Levels of 2 or 3 in the most recent NASA Center Disaster/Risk Matrix produced by the Climate Adaptation Science Investigator Working Group. Based on prior CASI workshops at other NASA centers, it is recognized that JSC is highly vulnerable to climate-change related hazards and has a need for adaptation strategies. We will present an overview of prior CASI-related work at JSC, including publication of a climate change and adaptation informational data brochure, and a Resilience and Adaptation to Climate Risks Workshop that was held at JSC in early March 2012. Major outcomes of that workshop that form a basis for work going forward are 1) a realization that JSC is embedded in a regional environmental and social context, and that potential climate change effects and adaptation strategies will not, and should not, be constrained by the Center fence line; 2) a desire to coordinate data collection and adaptation planning activities with interested stakeholders to form a regional climate change adaptation center that could facilitate interaction with CASI; 3) recognition that there is a wide array of basic data (remotely sensed, in situ, GIS/mapping, and historical) available through JSC and other stakeholders, but this data is not yet centrally accessible for planning purposes.

Stefanov, William L.↗

Introduction The Role of the Agricultural Model Intercomparison and Improvement Project

Climate impacts on agriculture are of increasing concern in both the scientific and policy communities because of the need to ensure food security for a growing population. A special challenge is posed by the changes in the frequency and intensity of heat-waves, droughts, and episodic rainstorms already underway in many parts of the world. Changes in production are directly linked to such variations in temperature and precipitation during the growing season, and often to offseason changes in weather affecting soil-water storage and availability to crops. This is not an isolated problem but one of both global and regional importance, because of impacts on the livelihoods of smallholder farmers as well as consequences for the world food trade system. This two-part set the Agricultural Model Intercomparison and Improvement Project (AgMIP): Integrated Crop and Economic Assessments is the first to be entirely devoted to AgMIP (www.agmip.org). AgMIP is a major international research program focused on climate change and agriculture. The goal of the two parts is to advance the field by providing detailed information on new simulation techniques and assessments being conducted by this program. It presents information about new methods of global and regional integrated assessment, results from agricultural regions, and adaptation strategies for maintaining food security under changing climate conditions.

security↗

Adaptive Management Methods to Protect the California Sacramento-San Joaquin Delta Water Resource

The California Sacramento-San Joaquin River Delta is the hub for California's water supply, conveying water from Northern to Southern California agriculture and communities while supporting important ecosystem services, agriculture, and communities in the Delta. Changes in climate, long-term drought, water quality changes, and expansion of invasive aquatic plants threatens ecosystems, impedes ecosystem restoration, and is economically, environmentally, and sociologically detrimental to the San Francisco Bay/California Delta complex. NASA Ames Research Center and the USDA-ARS partnered with the State of California and local governments to develop science-based, adaptive-management strategies for the Sacramento-San Joaquin Delta. The project combines science, operations, and economics related to integrated management scenarios for aquatic weeds to help land and waterway managers make science-informed decisions regarding management and outcomes. The team provides a comprehensive understanding of agricultural and urban land use in the Delta and the major water sheds (San Joaquin/Sacramento) supplying the Delta and interaction with drought and climate impacts on the environment, water quality, and weed growth. The team recommends conservation and modified land-use practices and aids local Delta stakeholders in developing management strategies. New remote sensing tools have been developed to enhance ability to assess conditions, inform decision support tools, and monitor management practices. Science gaps in understanding how native and invasive plants respond to altered environmental conditions are being filled and provide critical biological response parameters for Delta-SWAT simulation modeling. Operational agencies such as the California Department of Boating and Waterways provide testing and act as initial adopter of decision support tools. Methods developed by the project can become routine land and water management tools in complex river delta systems.

Agriculture↗

Acquisitive root exploration strategies help maintain higher peak sap flux rates during summer drought, but more root biomass does not

Roots are responsible for soil water uptake, yet little is known about how variation in fine-root traits relates to whole-tree water movement, particularly during periods of drought. By combining a 3-yr dataset monitoring sap flow rates with measures of fine-root biomass, length, and morphology across 10 tree species, we addressed hypotheses concerning the role of fine-root system size and morphology in determining tree responses to summer drought as well as potential changes in their relative importance under more severe droughts. Greater fine-root biomass and length did not enable trees to maintain high rates of sap flow during drought, whereas the morphological traits, specific root length and specific root area, were linked to sustained sap flow rates during drought. We found that all species, irrespective of root traits, progressively reduced their sap flow under more extreme drought conditions, although more acquisitive root morphology was still associated with smaller reductions. Our results run counter to long-standing assumptions that larger root systems are better able to access soil water and maintain photosynthetic activity during drought. Instead, we find evidence that root morphology at least partially determines the capacity for water uptake and movement as soil moisture declines.

drought↗

Data Fusion and Mining Techniques to Map Water Use and Drought across Spatial and Temporal Scales

As the world’s water resources come under increasing tension due to dual stressors of climate change and population growth, accurate knowledge of water consumption through evapotranspiration (ET) over a range in spatial scales will be critical in developing adaptation strategies. Remote sensing methods for monitoring consumptive water use (e.g, ET) are becoming increasingly important, especially in areas of significant water and food insecurity. One method to estimate ET from satellite-based methods, the Atmosphere Land Exchange Inverse (ALEXI) model uses the change in mid-morning land surface temperature to estimate the partitioning of sensible and latent heat fluxes which are then used to estimate daily ET. This presentation will outline several recent enhancements to the ALEXI modeling system, with a focus on global ET and drought monitoring. Until recently, ALEXI has been limited to areas with high resolution temporal sampling of geostationary sensors. The use of geostationary sensors makes global mapping a complicated process, especially for real-time applications, as data from as many as five different sensors are required to be ingested and harmonized to create a global mosaic. However, our research team has developed a new and novel method of using twice-daily observations from polar-orbiting sensors such as MODIS and VIIRS to estimate the mid-morning rise in LST that is used to drive the energy balance estimations within ALEXI. This allows the method to be applied globally using a single sensor (in this case, initially MODIS with a planned transition to VIIRS) rather than a global compositing of all available geostationary data. Other advantages of this new method include the higher spatial resolution provided by MODIS and VIIRS and the increased sampling at high latitudes where oblique view angles limit the utility of geostationary sensors. This presentation will focus on global applications for mapping water use and drought using data mining and data fusion across spatial scales extending from 5-km to 30-m “field-scale” estimates.

Christopher Hain↗

Watershed-Scale Modeling of Land-Use and Altered Environment Impacts on Aquatic Weed Growth in the Delta

The California Sacramento-San Joaquin River Delta is the hub for California's water supply, conveying water from Northern to Southern California agriculture and communities while supporting important ecosystem services, agriculture, and communities in the Delta. Changes in climate, long-term drought, and water quality have all been suspected as playing role in the dramatic expansion of invasive aquatic plants and their impact on ecosystems of the San Francisco Bay / California Delta complex. NASA Ames Research Center, USDA-Agricultural Research Service, the State of California, UC Davis, and local governments have partnered under a USDA sponsored project (DRAAWP) to develop science-based, adaptive-management strategies for invasive aquatic plants in Sacramento-San Joaquin Delta. Critical to developing management strategies is to understand how the Delta is affected by both the magnitude of fluctuations in land-use and climate / drought induced altered environments and how the plants respond to these altered environments. We utilize the Soil Water Assessment Tool (SWAT), a watershed-scale model developed to quantify the impact of land management practices in large and complex watersheds on water quality, as the backbone for a customized Delta model - Delta-SWAT. The model uses land-use, soils, elevation, and hydrologic routing to characterize pesticide and nutrient transport from the Sacramento and San Joaquin rivers watersheds and loading into the Delta. Land-use within the Delta, as well as water extraction to supply those functions, and the resulting return of water to Delta waterways are included in Delta-SWAT. Hydrologic transport within the Delta has required significant attention to address the lack of elevation driven transport processes. Delta-SWAT water quality trend estimates are compared with water quality monitoring conducted throughout the Delta. Aquatic plant response to water quality and other environmental factors is carried out using a customized model component. Plant response to the range of water quality factors, response times, and altered temperature and light regimes of the Delta have required gap-filling studies to provide model parameters. Delta-SWAT provides a tool for evaluating temporal and spatial effects of land-use and altered environments in the Delta and contributing watersheds on aquatic weed growth. Using Delta-SWAT for simulation modeling allows evaluation of historic and current conditions as well as consideration potential climate change and management practice outcomes. Delta-SWAT adds to the scientific understanding of dynamics in the Delta and enhances development of science-informed, management strategies and practices.

Ecosystem↗

The effects of environmental conditions on eggshell thickness of two North American secondary cavity-nesting birds

Egg hatchability in birds is a critical component of individual reproductive success and is associated with eggshell integrity. Contaminants, such as DDT, can influence eggshell thickness and are known to cause population declines. Moisture content and temperature can also impact eggshell thickness, but the influence of environmental conditions on the natural variation in eggshell thickness in wild populations is not well understood. Our goal was to investigate the relationship between environmental conditions and eggshell thickness of Western Bluebird (Sialia mexicana) and Ash-throated Flycatcher (Myiarchus cinerascens) eggs from 1997 to 2013 on the Pajarito Plateau in northern New Mexico. We tested whether nesting elevation, temperature, precipitation, and drought conditions influenced eggshell thickness in these two secondary cavity-nesting species, while also looking at eggshell thickness over time. Over the 16 years, nonviable or abandoned eggs were collected, and the analyzed dataset included 330 bluebird eggs and 113 flycatcher eggs. There was a significant increase in eggshell thickness over time for both species. Flycatcher eggshells were correlated with higher temperatures, whereas bluebird eggshells were influenced by an interaction between temperature and drought severity. In drought conditions, bluebird eggshell thickness was positively correlated with temperature, whereas in wet conditions, eggshell thickness was negatively correlated with temperature. Thicker eggshells in drought conditions may be a way of reducing water loss from eggs, which occurs faster at higher temperatures. In the southwestern United States, frequent and severe drought, higher temperatures, and decreases in precipitation are all expected to continue. In conclusion, the structure of eggs will be important to consider regarding how species may or may not adapt to novel conditions or persist in new environments.

59 BASIC BIOLOGICAL SCIENCES↗