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At least 235 records · Page 13

Solving the sample size problem for resource selection functions

Abstract Sample size sufficiency is a critical consideration for estimating resource selection functions (RSFs) from GPS‐based animal telemetry. Cited thresholds for sufficiency include a number of captured animals and as many relocations per animal N as possible. These thresholds render many RSF‐based studies misleading if large sample sizes were truly insufficient, or unpublishable if small sample sizes were sufficient but failed to meet reviewer expectations. We provide the first comprehensive solution for RSF sample size by deriving closed‐form mathematical expressions for the number of animals M and the number of relocations per animal N required for model outputs to a given degree of precision. The sample sizes needed depend on just 3 biologically meaningful quantities: habitat selection strength, variation in individual selection and a novel measure of landscape complexity, which we define rigorously. The mathematical expressions are calculable for any environmental dataset at any spatial scale and are applicable to any study involving resource selection (including sessile organisms). We validate our analytical solutions using globally relevant empirical data including 5,678,623 GPS locations from 511 animals from 10 species (omnivores, carnivores and herbivores living in boreal, temperate and tropical forests, montane woodlands, swamps and Arctic tundra). Our analytic expressions show that the required M and N must decline with increasing selection strength and increasing landscape complexity, and this decline is insensitive to the definition of availability used in the analysis. Our results demonstrate that the most biologically relevant effects on the utilization distribution (i.e. those landscape conditions with the greatest absolute magnitude of resource selection) can often be estimated with much fewer than animals. We identify several critical steps in implementing these equations, including (a) a priori selection of expected model coefficients and (b) regular sampling of background (pseudoabsence) data within a given definition of availability. We discuss possible methods to identify a priori expectations for habitat selection coefficients, effects of scale on RSF estimation and caveats for rare species applications. We argue that these equations should be a mandatory component for all future RSF studies.

Street, Garrett M.↗

Foliar functional traits from imaging spectroscopy across biomes in eastern North America

Summary Foliar functional traits are widely used to characterize leaf and canopy properties that drive ecosystem processes and to infer physiological processes in Earth system models. Imaging spectroscopy provides great potential to map foliar traits to characterize continuous functional variation and diversity, but few studies have demonstrated consistent methods for mapping multiple traits across biomes. With airborne imaging spectroscopy data and field data from 19 sites, we developed trait models using partial least squares regression, and mapped 26 foliar traits in seven NEON (National Ecological Observatory Network) ecoregions (domains) including temperate and subtropical forests and grasslands of eastern North America. Model validation accuracy varied among traits (normalized root mean squared error, 9.1–19.4%; coefficient of determination, 0.28–0.82), with phenolic concentration, leaf mass per area and equivalent water thickness performing best across domains. Across all trait maps, 90% of vegetated pixels had reasonable values for one trait, and 28–81% provided high confidence for multiple traits concurrently. Maps of 26 traits and their uncertainties for eastern US NEON sites are available for download, and are being expanded to the western United States and tundra/boreal zone. These data enable better understanding of trait variations and relationships over large areas, calibration of ecosystem models, and assessment of continental‐scale functional diversity.

Wang, Zhihui↗

Model-ready benchmarks for NPP, ANPP, litter fluxes, and recruitment into the 1 cm dbh size class

The intended use of this dataset is to serve as an observational benchmark to evaluate model predictions of NPP, ANPP, litter fluxes, and recruitment at Barro Colorado Island, Panama. This dataset contains four CSV files and one text file. “Benchmarks-NPP-ANPP-R-L.csv” provides estimates of annual ecosystem-level reproductive litter flux (R), leaf litter flux (L), aboveground net primary productivity (ANPP), and net primary productivity (NPP) for 61 field plots throughout tropical, temperate, and boreal forest biomes. An additional 499 plots (n = 550) include estimates of just R, L, and R/L. Each row reports a distinct set of estimates for one sampling interval at one plot. “Metadata-Benchmarks-NPP-ANPP-R-L.csv” contains field descriptions for all data fields in “Benchmarks-NPP-ANPP-R-L.csv”. “References-Benchmarks-NPP-ANPP-R-L.txt” contains full references to the original studies used to produce the observations at each plot included in the data. “Benchmarks-Recruitment.csv” provides estimates of species-level recruitment rates into the 1 cm dbh size class at four CTFS-ForestGeo sites using methods that account for unobserved mortality of new recruits between census intervals (Kohyama et al., 2018). “Metadata-Benchmarks-Recruitment.csv” contains field descriptions for all data fields in “Benchmarks-Recruitment.csv”.

54 ENVIRONMENTAL SCIENCES↗

Global 10 m Land Use Land Cover Datasets: A Comparison of Dynamic World, World Cover and Esri Land Cover

The European Space Agency’s Sentinel satellites have laid the foundation for global land use land cover (LULC) mapping with unprecedented detail at 10 m resolution. We present a cross-comparison and accuracy assessment of Google’s Dynamic World (DW), ESA’s World Cover (WC) and Esri’s Land Cover (Esri) products for the first time in order to inform the adoption and application of these maps going forward. For the year 2020, the three global LULC maps show strong spatial correspondence (i.e., near-equal area estimates) for water, built area, trees and crop LULC classes. However, relative to one another, WC is biased towards over-estimating grass cover, Esri towards shrub and scrub cover and DW towards snow and ice. Using global ground truth data with a minimum mapping unit of 250 m 2 , we found that Esri had the highest overall accuracy (75%) compared to DW (72%) and WC (65%). Across all global maps, water was the most accurately mapped class (92%), followed by built area (83%), tree cover (81%) and crops (78%), particularly in biomes characterized by temperate and boreal forests. The classes with the lowest accuracies, particularly in the tundra biome, included shrub and scrub (47%), grass (34%), bare ground (57%) and flooded vegetation (53%). When using European ground truth data from LUCAS (Land Use/Cover Area Frame Survey) with a minimum mapping unit of <100 m 2 , we found that WC had the highest accuracy (71%) compared to DW (66%) and Esri (63%), highlighting the ability of WC to resolve landscape elements with more detail compared to DW and Esri. Although not analyzed in our study, we discuss the relative advantages of DW due to its frequent and near real-time data delivery of both categorical predictions and class probability scores. We recommend that the use of global LULC products should involve critical evaluation of their suitability with respect to the application purpose, such as aggregate changes in ecosystem accounting versus site-specific change detection in monitoring, considering trade-offs between thematic resolution, global versus. local accuracy, class-specific biases and whether change analysis is necessary. We also emphasize the importance of not estimating areas from pixel-counting alone but adopting best practices in design-based inference and area estimation that quantify uncertainty for a given study area.

54 ENVIRONMENTAL SCIENCES↗

Hyperspectral Remote Sensing of Foliar Nitrogen Content

A strong positive correlation between vegetation canopy bidirectional reflectance factor (BRF) in the near infrared (NIR) spectral region and foliar mass-based nitrogen concentration (%N) has been reported in some temperate and boreal forests. This relationship, if true, would indicate an additional role for nitrogen in the climate system via its influence on surface albedo and may offer a simple approach for monitoring foliar nitrogen using satellite data. We report, however, that the previously reported correlation is an artifact - it is a consequence of variations in canopy structure, rather than of %N. The data underlying this relationship were collected at sites with varying proportions of foliar nitrogen-poor needleleaf and nitrogen-rich broadleaf species, whose canopy structure differs considerably. When the BRF data are corrected for canopy-structure effects, the residual reflectance variations are negatively related to %N at all wavelengths in the interval 423-855 nm. This suggests that the observed positive correlation between BRF and %N conveys no information about %N. We find that to infer leaf biochemical constituents, e.g., N content, from remotely sensed data, BRF spectra in the interval 710-790 nm provide critical information for correction of structural influences. Our analysis also suggests that surface characteristics of leaves impact remote sensing of its internal constituents. This further decreases the ability to remotely sense canopy foliar nitrogen. Finally, the analysis presented here is generic to the problem of remote sensing of leaf-tissue constituents and is therefore not a specific critique of articles espousing remote sensing of foliar %N.

Knyazikhin, Yuri↗

Variability of Phenology and Fluxes of Water and Carbon with Observed and Simulated Soil Moisture in the Ent Terrestrial Biosphere Model (Ent TBM Version 1.0.1.0.0)

The Ent Terrestrial Biosphere Model (Ent TBM) is a mixed-canopy dynamic global vegetation model developed specifically for coupling with land surface hydrology and general circulation models (GCMs). This study describes the leaf phenology submodel implemented in the Ent TBM version 1.0.1.0.0 coupled to the carbon allocation scheme of the Ecosystem Demography (ED) model. The phenology submodel adopts a combination of responses to temperature (growing degree days and frost hardening), soil moisture (linearity of stress with relative saturation) and radiation (light length). Growth of leaves, sapwood, fine roots, stem wood and coarse roots is updated on a daily basis. We evaluate the performance in reproducing observed leaf seasonal growth as well as water and carbon fluxes for four plant functional types at five Fluxnet sites, with both observed and prognostic hydrology, and observed and prognostic seasonal leaf area index. The phenology submodel is able to capture the timing and magnitude of leaf-out and senescence for temperate broadleaf deciduous forest (Harvard Forest and Morgan- Monroe State Forest, US), C3 annual grassland (Vaira Ranch, US) and California oak savanna (Tonzi Ranch, US). For evergreen needleleaf forest (Hyytiäla, Finland), the phenology submodel captures the effect of frost hardening of photosynthetic capacity on seasonal fluxes and leaf area. We address the importance of customizing parameter sets of vegetation soil moisture stress response to the particular land surface hydrology scheme. We identify model deficiencies that reveal important dynamics and parameter needs.

Soil moisture↗

Substrate quality drives fungal necromass decay and decomposer community structure under contrasting vegetation types

Fungal mycelium is increasingly recognized as a central component of soil biogeochemical cycling, yet our current understanding of the ecological controls on fungal necromass decomposition is limited to single sites and vegetation types. By deploying common fungal necromass substrates in a temperate oak savanna and hardwood forest in the midwestern USA, we assessed the generality of the rate at which high- and low-quality fungal necromass decomposes; further, we investigated how the decomposer ‘necrobiome’ varies both across and within sites under vegetation types dominated by either arbuscular or ectomycorrhizal plants. In this work, the effects of necromass quality on decay rate were robust to site and vegetation type differences, with high-quality fungal necromass decomposing, on average, 2.5 times faster during the initial stages of decay. Across vegetation types, bacterial and fungal communities present on decaying necromass differed from bulk soil microbial communities and were influenced by necromass quality. Moulds, yeasts and copiotrophic bacteria consistently dominated the necrobiome of high-quality fungal substrates. Synthesis. We show that regardless of differences in decay environments, high-quality fungal substrates decompose faster and support different types of decomposer micro-organisms when compared with low-quality fungal tissues. These findings help to refine our theoretical understanding of the dominant factors affecting fast cycling components of soil organic matter and the microbial communities associated with rapid decay.

59 BASIC BIOLOGICAL SCIENCES↗

Linking leaf dark respiration to leaf traits and reflectance spectroscopy across diverse forest types

Leaf dark respiration (R dark ), an important yet rarely quantified component of carbon cycling in forest ecosystems, is often simulated from leaf traits such as the maximum carboxylation capacity (V cmax ), leaf mass per area (LMA), nitrogen (N) and phosphorus (P) concentrations, in terrestrial biosphere models. However, the validity of these relationships across forest types remains to be thoroughly assessed. Here, in this study, we analyzed R dark variability and its associations with V cmax and other leaf traits across three temperate, subtropical and tropical forests in China, evaluating the effectiveness of leaf spectroscopy as a superior monitoring alternative. We found that leaf magnesium and calcium concentrations were more significant in explaining cross-site R dark than commonly used traits like LMA, N and P concentrations, but univariate trait–R dark relationships were always weak (r 2 ≤ 0.15) and forest-specific. Although multivariate relationships of leaf traits improved the model performance, leaf spectroscopy outperformed trait–R dark relationships, accurately predicted cross-site R dark (r 2 = 0.65) and pinpointed the factors contributing to R dark variability. Our findings reveal a few novel traits with greater cross-site scalability regarding R dark , challenging the use of empirical trait–R dark relationships in process models and emphasize the potential of leaf spectroscopy as a promising alternative for estimating R dark , which could ultimately improve process modeling of terrestrial plant respiration.

59 BASIC BIOLOGICAL SCIENCES↗

Mycorrhizal associations of tree species influence soil nitrogen dynamics via effects on soil acid–base chemistry

Across all spatial scales, ECM-dominated forests were associated with greater soil acidity. In particular, ECM-dominated soils exhibited lower soil pH and base cations, although the magnitude of mycorrhizal-associated differences in soil acid–base chemistry depended on the biomes, with differences being more pronounced in temperate than in sub/tropical forests. Higher lignin and lower base cations in ECM tree leaf litter were related to greater soil acidity in ECM-dominated forests. Moreover, the lower inorganic N concentrations and slower N transformation rates in ECM-dominated forests were associated with their greater soil acidity. Our results indicate that the scale-invariant feedbacks between plant nutrient-use strategies and soil properties have the potential to impact forest community assembly and ecosystem processes, particularly in the context of global change.

37 INORGANIC, ORGANIC, PHYSICAL, AND ANALYTICAL CH↗

Forest aboveground biomass estimation through integration of sentinel-2 and PALSAR-2 time series: assessing models trained on GEDI and field inventory benchmarks

Accurate and spatially explicit forest Aboveground Biomass (AGB) mapping through remote sensing is critical for quantifying terrestrial carbon stocks and informing effective forest management strategies. However, AGB estimation in dense forests with complex terrain remains challenging due to satellite sensor signal saturation problem (saturation issue occurs in high biomass forests), structural complexity, and limited ground truth for calibration. This study presents a novel framework that integrates multi-temporal Sentinel-2 optical imagery, ALOS PALSAR-2 Synthetic Aperture Radar (SAR) data, and topographic variables with explainable Machine Learning to map AGB across mountainous forests within subtropical and temperate oceanic climate zones of Mexico. We evaluate the effects of temporal granularity and sensor synergy by comparing multiple temporal inputs and sensor configurations (Sentinel-2, PALSAR-2, and their fusion), and assess model performance using two reference datasets: NASA GEDI LiDAR-derived biomass and Mexico’s National Forest and Soil Inventory (INFyS). Our results showed that models trained on INFyS consistently outperformed those trained on GEDI, highlighting limitations in GEDI’s reliability in biomass estimates within this study region. Furthermore, the integration of Sentinel-2 and PALSAR-2 provided improved predictions compared to single-sensor models, particularly when combined with temporally explicit yearly statistics. The best-performing model, which was trained on INFyS data, and considered both Sentinel-2 and PALSAR-2 yearly statistics, as well as topographic variables, achieved an R2 of 0.64, RMSE of 51.10 Mg/ha, and relative RMSE (rRMSE) of 58.69%. Explainable ML analysis identified Sentinel-2 spectral indices and topographic features as key predictors, while PALSAR-2 metrics provided complementary information, partially mitigating saturation effects in high-biomass areas. Specifically, integrating both sensors substantially improved AGB estimation in high biomass forest (≥200 Mg/ha), yielding 98% gains over optical-only model, with resulting estimates exceeding GEDI L4B by 29% and ESA-CCI-BIOMASS by 174%. Terrain-stratified analysis indicated close agreement with GEDI in low-slope areas, with increasing divergence as slope steepness increased, while estimates remained consistently higher than ESA-CCI-BIOMASS across all slope classes. The proposed approach advances multi-sensor fusion and temporal feature engineering for AGB mapping using open-access satellite datasets, providing a scalable and reproducible framework for annual biomass monitoring in topographically complex mountainous forests. The resulting 25 m resolution biomass product has the potential to provide spatially detailed information for forest monitoring and may support applications in carbon accounting and forest management.

54 ENVIRONMENTAL SCIENCES↗

Tree drought physiology: critical research questions and strategies for mitigating climate change effects on forests

Droughts of increasing severity and frequency are a primary cause of forest mortality associated with climate change. Yet, fundamental knowledge gaps regarding the complex physiology of trees limit the development of more effective management strategies to mitigate drought effects on forests. Here, in this work, we highlight some of the basic research needed to better understand tree drought physiology and how new technologies and interdisciplinary approaches can be used to address them. Our discussion focuses on how trees change wood development to mitigate water stress, hormonal responses to drought, genetic variation underlying adaptive drought phenotypes, how trees ‘remember’ prior stress exposure, and how symbiotic soil microbes affect drought response. Next, we identify opportunities for using research findings to enhance or develop new strategies for managing drought effects on forests, ranging from matching genotypes to environments, to enhancing seedling resilience through nursery treatments, to landscape-scale monitoring and predictions. We conclude with a discussion of the need for co-producing research with land managers and extending research to forests in critical ecological regions beyond the temperate zone.

54 ENVIRONMENTAL SCIENCES↗

Radar detection of flooding beneath the forest canopy - A review

Synthetic aperture radar remote sensing is a promising tool for detection of flooding on forested floodplains. The brigtht appearance of flooded forests on radar images results from double-bounce reflections between smooth water surfaces and tree trunks or branches. Enhanced backscattering at L-band has been shown to occur in a wide variety of forest types, including cypress-tupelo swamps, temperate bottomland hardwoods, spruce bogs, mangroves and tropical floodplain forests. Lack of enhancement is a function of both stand density and branching structure. According to models and measurements, the magnitude of the enhancement is about 3 to 10 dB. Steep incidence angles (20-30 deg) are optimal for detection of flooding, since some forest types exhibit bright returns only at steeper angles. P-band should prove useful for floodwater mapping in dense stands, and multifrequency polarimetric analysis should allow flooded forests to be distinguished from marshes.

Hess, Laura L.↗

Ectomycorrhizal effects on decomposition are highly dependent on fungal traits, climate, and litter properties: A model-based assessment

It has been proposed that competition between ectomycorrhizal (ECM) fungi and free-living saprotrophs for resources like nitrogen (N) slows decomposition and increases the soil carbon storage in ECM ecosystems compared to arbuscular (AM) ecosystems. However, empirical evidence for the generality of such ECM effects is equivocal, and confounding mechanisms have been proposed that affect the magnitude and direction of ECM effects on soil carbon. Here, for this work, we conduct a theoretical modeling experiment, where we explicitly incorporate mycorrhizal processes into the Carbon, Organisms, Rhizosphere, and Protection in the Soil Environment (CORPSE) model. We use the model to explore the conditions under which ECM N acquisition processes can induce stronger saprotrophic N limitation and result in slower decomposition rates and greater soil organic carbon accumulation compared to AM processes. We found that the ECM fungi more strongly inhibited decomposition when litter inputs were N-depleted and relatively recalcitrant and when ECM fungi possessed a strong capacity to mine N from both recalcitrant soil organic matter and microbial necromass. Climate and seasonality also played a role as the ECM competition effect was strongest at low mean annual temperatures and when litterfall peaked seasonally. Priming effects driven by high root exudation rates in ECM-dominated systems could overwhelm the competition effect and reduce soil carbon under some circumstances. The ECM effect on decomposition in our simulations was highly context dependent. Based on our model results, we expect to see a strong ECM competition effect in temperate deciduous and boreal forests with relatively recalcitrant litter inputs, and with ECM fungi that produce oxidases and necromass-degrading enzymes. However, even a relatively strong ECM competition effect on decomposition only increased soil organic carbon accumulation by ~10%.

59 BASIC BIOLOGICAL SCIENCES↗

Characterizing the Atmospheric Mn Cycle and Its Impact on Terrestrial Biogeochemistry

Abstract The role of manganese (Mn) in ecosystem carbon (C) biogeochemical cycling is gaining increasing attention. While soil Mn is mainly derived from bedrock, atmospheric deposition could be a major source of Mn to surface soils, with implications for soil C cycling. However, quantification of the atmospheric Mn cycle, which comprises emissions from natural (desert dust, sea salts, volcanoes, primary biogenic particles, and wildfires) and anthropogenic sources (e.g., industrialization and land‐use change due to agriculture), transport, and deposition, remains uncertain. Here, we use compiled emission data sets for each identified source to model and quantify the atmospheric Mn cycle by combining an atmospheric model and in situ atmospheric concentration measurements. We estimated global emissions of atmospheric Mn in aerosols (<10 μm in aerodynamic diameter) to be 1,400 Gg Mn year −1 . Approximately 31% of the emissions come from anthropogenic sources. Deposition of the anthropogenic Mn shortened Mn “pseudo” turnover times in 1‐m‐thick surface soils (ranging from 1,000 to over 10,000,000 years) by 1–2 orders of magnitude in industrialized regions. Such anthropogenic Mn inputs boosted the Mn‐to‐N ratio of the atmospheric deposition in non‐desert dominated regions (between 5 × 10 −5 and 0.02) across industrialized areas, but that was still lower than soil Mn‐to‐N ratio by 1–3 orders of magnitude. Correlation analysis revealed a negative relationship between Mn deposition and topsoil C density across temperate and (sub)tropical forests, consisting with atmospheric Mn deposition enhancing carbon respiration as seen in in situ biogeochemical studies.

Environmental Sciences & Ecology↗

Characterization of terrestrial ecosystems for biogeochemical studies using remote sensing

Work is in progress to estimate leaf area index (LAI) of temperate closed canopy coniferous forests using transects in Oregon and California. This variable will be measured using remote sensing techniques including correlations of ground dimensional analysis with linear waveband combinations. LAI will be related to important biological variables such as net primary productivity, biomass, and biogenic gas emission fluxes. The spatial variation in LAI, when coupled with species composition, will be used in part to describe the spatial variation and temporal dynamics of biogeochemical cycling.

Peterson, D. L.↗

Calibrated imaging radar polarimetry - Technique, examples, and applications

The authors developed a calibration procedure for imaging radar polarimeters and applied it to a set of images acquired by the NASA DC-8 multifrequency radar system. The technique requires the use of ground reflectors known cross-section for absolute calibration, that is, solution for sigma exp 0; however, the image data themselves can usually provide all information necessary for phase calibration and for antenna crosstalk correction. The accuracy of the approach, as measured by calculating the cross-section residuals of known targets in each calibrated scene, is on the order of +/- 1-2 dB at P- and C-band, but improves to +/- 0.5 dB at L-band. The authors present the results of applying this technique to radar scenes of lava flows of varying roughness, temperate and tropical rain forests, and ocean water surfaces. They also present several example applications which are feasible with calibrated data but which would be difficult to implement with uncalibrated data.

Zebker, Howard A.↗

The Net Exchange Between Terrestrial Ecosystems and the Atmosphere as a Result of Changes in Land Use

The general purpose of this research was to improve and update (to 1990) estimates of the net flux of carbon between the world's terrestrial ecosystems and the atmosphere from changes in land use (e.g., deforestation and reforestation). The estimates are important for understanding the global carbon cycle, and for predicting future concentrations of atmospheric CO2 that will result from emissions. The emphasis of the first year's research was on the northern temperate zone and boreal forests, where the greatest discrepancy exists between estimates of flux. Forest inventories suggest net sinks of 0.6 PgC/yr; inversion analyses based on atmospheric data and models suggest much larger sinks 2-3.6 PgC/yr (e.g., Tans et al. 1990, Ciais et al. 1995). The work carried out with this grant calculated the flux attributable to changes in land use. The estimated flux was somewhat smaller than the flux calculated from inventory data suggesting that environmental changes have led to a small accumulation of carbon in forests that exceeds the accumulation expected from past rates of harvest. Two publications have described these results (Houghton 1996, 1998). The large difference between these estimates and those obtained with atmospheric data and models remains unexplained. The recent estimate of a 1.7 PgC/yr sink in North America, alone (Fan et al. 1998), is particularly difficult to explain. That part of the sink attributable to land-use change, however, is defined as a result of this grant.

Houghton, R. A.↗

Remote Sensing of the Urban Heat Island Effect Across Biomes in the Continental USA

Impervious surface area (ISA) from the Landsat TM-based NLCD 2001 dataset and land surface temperature (LST) from MODIS averaged over three annual cycles (2003-2005) are used in a spatial analysis to assess the urban heat island (UHI) skin temperature amplitude and its relationship to development intensity, size, and ecological setting for 38 of the most populous cities in the continental United States. Development intensity zones based on %ISA are defined for each urban area emanating outward from the urban core to the nonurban rural areas nearby and used to stratify sampling for land surface temperatures and NDVI. Sampling is further constrained by biome and elevation to insure objective intercomparisons between zones and between cities in different biomes permitting the definition of hierarchically ordered zones that are consistent across urban areas in different ecological setting and across scales. We find that ecological context significantly influences the amplitude of summer daytime UHI (urban-rural temperature difference) the largest (8 C average) observed for cities built in biomes dominated by temperate broadleaf and mixed forest. For all cities combined, ISA is the primary driver for increase in temperature explaining 70% of the total variance in LST. On a yearly average, urban areas are substantially warmer than the non-urban fringe by 2.9 C, except for urban areas in biomes with arid and semiarid climates. The average amplitude of the UHI is remarkably asymmetric with a 4.3 C temperature difference in summer and only 1.3 C in winter. In desert environments, the LST's response to ISA presents an uncharacteristic "U-shaped" horizontal gradient decreasing from the urban core to the outskirts of the city and then increasing again in the suburban to the rural zones. UHI's calculated for these cities point to a possible heat sink effect. These observational results show that the urban heat island amplitude both increases with city size and is seasonally asymmetric for a large number of cities across most biomes. The implications are that for urban areas developed within forested ecosystems the summertime UHI can be quite high relative to the wintertime UHI suggesting that the residential energy consumption required for summer cooling is likely to increase with urban growth within those biomes.

Imhoff, Marc L.↗